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Genetic study Ancient genomes reveal trans-Eurasian connections between the European Huns and the Xiongnu Empire

Tautalus

Regular Member
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Portuguese
Y-DNA haplogroup
I2-M223 / I-FTB15368
mtDNA haplogroup
H6a1b2y
This paper examines the genetic connections between the European Huns and the Xiongnu Empire by analyzing genomic data from 370 individuals across different periods and regions. The findings indicate that the Huns, who appeared in Europe in the 4th century, might have genetic ties to the Xiongnu Empire, which dissolved around 300 years before. Despite high genetic diversity among Hun-period burials in the Carpathian Basin, there is evidence of a mixed origin of steppe conquerors. Some European Huns descended from Xiongnu-period elites, suggesting trans-Eurasian genetic connections. The research provides new insights into the origins and diversity of the Hun-period population, showing long-shared genomic tracts connecting individuals from the highest Xiongnu-period elites to those in the Carpathian Basin. The study highlights the mixed ancestry of the incoming steppe conquerors and the persistence of some Hunnic cultural elements in the region. These findings allow us to better understand the genetic and cultural links between the European Huns and their ancestors in the Eurasian steppes.
Abstract
The Huns appeared in Europe in the 370s, establishing an Empire that reshaped West Eurasian history. Yet until today their origins remain a matter of extensive debate. Traditional theories link them to the Xiongnu, the founders of the first nomadic empire of the Mongolian steppe. The Xiongnu empire dissolved, however, ~300 y before the Huns appeared in Europe, and there is little archaeological and historical evidence of Huns in the steppe during this time gap. Furthermore, despite the rich 5th to 6th centuries current era (CE) archaeological record of the Carpathian Basin, the cultural elements of connections with the steppe are limited to few findings and even fewer solitary eastern-type burials. In this study, we coanalyze archaeological evidence with 35 newly sequenced and published genomic data for a total of 370 individuals—from 5th to 6th century CE contexts in the Carpathian Basin including 10 Hun-period eastern-type burials, 2nd to 5th century sites across Central Asia and 2nd c. before current era (BCE) to 1st c. CE Xiongnu period sites across the Mongolian steppe. We find no evidence for the presence of a large eastern/steppe descent community among the Hun and post-Hun-period Carpathian Basin population. We also observe a high genetic diversity among the eastern-type burials that recapitulates the variability observed across the Eurasian Steppe. This suggests a mixed origin of the incoming steppe conquerors. Nevertheless, long-shared genomic tracts provide compelling evidence of genetic lineages directly connecting some individuals of the highest Xiongnu-period elite with 5th to 6th century CE Carpathian Basin individuals, showing that some European Huns descended from them.

Ancient genomes reveal trans-Eurasian connections between the European Huns and the Xiongnu Empire

PCA and Best qpAdm models (P-value > 0.01) of the Carpathian Basin individuals from the 10 Hun period eastern-type-burials (Top) and from the Hun or Gepid period (5th to 6th c. CE) context presenting East Eurasian/Steppe admixture signals.
s6ibT48.jpg


Haplogroups of the 35 newly sequenced individuals.

Genetic_IDGroupsexyhapmtHap
BRE024Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEMR1a1a(R-M512)U5a1d2b
BRE026Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEMR1(R-M306,R-M173)F1b1b
BRE018Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEMQ1a2a(Q-L475,Q-L53)C4a1a+195
BRE020Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEMQ1a1b1a(Q-L715,Q-L713)B4b1a3a
BRE031_ssBerel_Altai_Xianbei_Hun_P 2nd-5th c. CEMO1b1a1a1b(O-F3092,O-M1283)C4a1a3
BRE021Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEFA8a
BRE017Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEFC4a1a4
BRE022Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEFD4j+(16286)
BRE019Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEFH7b
BRE025Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEFH7b
BRE023Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEFU1a1a1
BRE030_ssBerel_Altai_Xianbei_Hun_P 2nd-5th c. CEFU5a1d2b
HDB001.A0101.ssModHun P eastern-type-burials 5th c. CEMR1a1a1b2a(R-Z94)U4b1a4
NEP2Hun P eastern-type-burials 5th c. CEMG2a1a1a2(G-FGC1159,G-FGC1160)U5a1d1
NEP1Hun P eastern-type-burials 5th c. CEME1b1b1a1b1(E-CTS1773,E-L618)H
PTL013.A0101.ssModHun P eastern-type-burials 5th c. CEFC4a1
TGH028.A0101.ssModHun/Gepid P 5th - 6th c. CEMR1b1(R-M415,R-P25_1)H6a1a
TGB023.A0101.ssModHun/Gepid P 5th - 6th c. CEMR1a1a1b2a(R-Z94)U5a2b1c
TGH010.A0101.ssModHun/Gepid P 5th - 6th c. CEMR1(R-M306,R-M173)D4o1
TGH012.A0101.ssModHun/Gepid P 5th - 6th c. CEMN1c1a1(N-L708)T1a1
TGH058.A0101.ssModHun/Gepid P 5th - 6th c. CEFC4a2c1
PTL004.A0101.ssModHun/Gepid P 5th - 6th c. CEFH1a
TGB005.A0101.ssModHun/Gepid P 5th - 6th c. CEFH1c
PTL002.A0101.ssModHun/Gepid P 5th - 6th c. CEFH26
PTL003.A0101.ssModHun/Gepid P 5th - 6th c. CEFH26
TGH019.A0101.ssModHun/Gepid P 5th - 6th c. CEFH80
TGH068.A0101.ssModHun/Gepid P 5th - 6th c. CEFJ1c5a
TGH016.A0101.ssModHun/Gepid P 5th - 6th c. CEFU3b
TGH015.A0101.ssModHun/Gepid P 5th - 6th c. CEFU5a2b1c
TGB013.A0101.ssModlateSarmatian_Hun_P 4th-5th c. CEMR1a(R-L62,R-M420)H3p
HNF075.A0101.ssModlateSarmatian_Hun_P 4th-5th c. CEMQ1a1b1(Q-L712)J1
HNF054.A0101.ssModlateSarmatian_Hun_P 4th-5th c. CEMI2a2a1a2a1a1(I-Z2054)U4
HNF016.A0101.ssModlateSarmatian_Hun_P 4th-5th c. CEMI2a2a1a2a(I-L1229)U5a2b
HNF019.A0101.ssModlateSarmatian_Hun_P 4th-5th c. CEMD1b2a(D-CTS6287,D-CTS220)H4a1a4b
HNF052.A0101.ssModlateSarmatian_Hun_P 4th-5th c. CEFU4b



 
This paper examines the genetic connections between the European Huns and the Xiongnu Empire by analyzing genomic data from 370 individuals across different periods and regions. The findings indicate that the Huns, who appeared in Europe in the 4th century, might have genetic ties to the Xiongnu Empire, which dissolved around 300 years before. Despite high genetic diversity among Hun-period burials in the Carpathian Basin, there is evidence of a mixed origin of steppe conquerors. Some European Huns descended from Xiongnu-period elites, suggesting trans-Eurasian genetic connections. The research provides new insights into the origins and diversity of the Hun-period population, showing long-shared genomic tracts connecting individuals from the highest Xiongnu-period elites to those in the Carpathian Basin. The study highlights the mixed ancestry of the incoming steppe conquerors and the persistence of some Hunnic cultural elements in the region. These findings allow us to better understand the genetic and cultural links between the European Huns and their ancestors in the Eurasian steppes.
Abstract
The Huns appeared in Europe in the 370s, establishing an Empire that reshaped West Eurasian history. Yet until today their origins remain a matter of extensive debate. Traditional theories link them to the Xiongnu, the founders of the first nomadic empire of the Mongolian steppe. The Xiongnu empire dissolved, however, ~300 y before the Huns appeared in Europe, and there is little archaeological and historical evidence of Huns in the steppe during this time gap. Furthermore, despite the rich 5th to 6th centuries current era (CE) archaeological record of the Carpathian Basin, the cultural elements of connections with the steppe are limited to few findings and even fewer solitary eastern-type burials. In this study, we coanalyze archaeological evidence with 35 newly sequenced and published genomic data for a total of 370 individuals—from 5th to 6th century CE contexts in the Carpathian Basin including 10 Hun-period eastern-type burials, 2nd to 5th century sites across Central Asia and 2nd c. before current era (BCE) to 1st c. CE Xiongnu period sites across the Mongolian steppe. We find no evidence for the presence of a large eastern/steppe descent community among the Hun and post-Hun-period Carpathian Basin population. We also observe a high genetic diversity among the eastern-type burials that recapitulates the variability observed across the Eurasian Steppe. This suggests a mixed origin of the incoming steppe conquerors. Nevertheless, long-shared genomic tracts provide compelling evidence of genetic lineages directly connecting some individuals of the highest Xiongnu-period elite with 5th to 6th century CE Carpathian Basin individuals, showing that some European Huns descended from them.

Ancient genomes reveal trans-Eurasian connections between the European Huns and the Xiongnu Empire

PCA and Best qpAdm models (P-value > 0.01) of the Carpathian Basin individuals from the 10 Hun period eastern-type-burials (Top) and from the Hun or Gepid period (5th to 6th c. CE) context presenting East Eurasian/Steppe admixture signals.
s6ibT48.jpg


Haplogroups of the 35 newly sequenced individuals.

Genetic_IDGroupsexyhapmtHap
BRE024Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEMR1a1a(R-M512)U5a1d2b
BRE026Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEMR1(R-M306,R-M173)F1b1b
BRE018Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEMQ1a2a(Q-L475,Q-L53)C4a1a+195
BRE020Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEMQ1a1b1a(Q-L715,Q-L713)B4b1a3a
BRE031_ssBerel_Altai_Xianbei_Hun_P 2nd-5th c. CEMO1b1a1a1b(O-F3092,O-M1283)C4a1a3
BRE021Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEFA8a
BRE017Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEFC4a1a4
BRE022Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEFD4j+(16286)
BRE019Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEFH7b
BRE025Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEFH7b
BRE023Berel_Altai_Xianbei_Hun_P 2nd-5th c. CEFU1a1a1
BRE030_ssBerel_Altai_Xianbei_Hun_P 2nd-5th c. CEFU5a1d2b
HDB001.A0101.ssModHun P eastern-type-burials 5th c. CEMR1a1a1b2a(R-Z94)U4b1a4
NEP2Hun P eastern-type-burials 5th c. CEMG2a1a1a2(G-FGC1159,G-FGC1160)U5a1d1
NEP1Hun P eastern-type-burials 5th c. CEME1b1b1a1b1(E-CTS1773,E-L618)H
PTL013.A0101.ssModHun P eastern-type-burials 5th c. CEFC4a1
TGH028.A0101.ssModHun/Gepid P 5th - 6th c. CEMR1b1(R-M415,R-P25_1)H6a1a
TGB023.A0101.ssModHun/Gepid P 5th - 6th c. CEMR1a1a1b2a(R-Z94)U5a2b1c
TGH010.A0101.ssModHun/Gepid P 5th - 6th c. CEMR1(R-M306,R-M173)D4o1
TGH012.A0101.ssModHun/Gepid P 5th - 6th c. CEMN1c1a1(N-L708)T1a1
TGH058.A0101.ssModHun/Gepid P 5th - 6th c. CEFC4a2c1
PTL004.A0101.ssModHun/Gepid P 5th - 6th c. CEFH1a
TGB005.A0101.ssModHun/Gepid P 5th - 6th c. CEFH1c
PTL002.A0101.ssModHun/Gepid P 5th - 6th c. CEFH26
PTL003.A0101.ssModHun/Gepid P 5th - 6th c. CEFH26
TGH019.A0101.ssModHun/Gepid P 5th - 6th c. CEFH80
TGH068.A0101.ssModHun/Gepid P 5th - 6th c. CEFJ1c5a
TGH016.A0101.ssModHun/Gepid P 5th - 6th c. CEFU3b
TGH015.A0101.ssModHun/Gepid P 5th - 6th c. CEFU5a2b1c
TGB013.A0101.ssModlateSarmatian_Hun_P 4th-5th c. CEMR1a(R-L62,R-M420)H3p
HNF075.A0101.ssModlateSarmatian_Hun_P 4th-5th c. CEMQ1a1b1(Q-L712)J1
HNF054.A0101.ssModlateSarmatian_Hun_P 4th-5th c. CEMI2a2a1a2a1a1(I-Z2054)U4
HNF016.A0101.ssModlateSarmatian_Hun_P 4th-5th c. CEMI2a2a1a2a(I-L1229)U5a2b
HNF019.A0101.ssModlateSarmatian_Hun_P 4th-5th c. CEMD1b2a(D-CTS6287,D-CTS220)H4a1a4b
HNF052.A0101.ssModlateSarmatian_Hun_P 4th-5th c. CEFU4b


thanks for sharing ;)
the D case was to good to be true
infact individual HNF019 fall under y haplogroup I
i checked him with the ytree site

 
It is either a wrong y-haplogroup assignment made by the authors in the paper or a wrong assignment made on the Ytree website. Does Ytree just record the information present in the paper or does it reanalyze the haplogroups ?
In the paper HNF016 is I-L1229, if it is related to HNF019 it is logical that they have the same y-haplogroup.
 
It is either a wrong y-haplogroup assignment made by the authors in the paper or a wrong assignment made on the Ytree website. Does Ytree just record the information present in the paper or does it reanalyze the haplogroups ?
In the paper HNF016 is I-L1229, if it is related to HNF019 it is logical that they have the same y-haplogroup.

I think they reanalyzed
I was excited to see D case
So to be sure i ask them to look at him 🤔
They gave me this link
To samples in theytree from that specific paper:

 
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